At the western shores of Messinian Mani Peninsula in South Greece, the composite, integrated karstic system of ‘‘Selinitsa’’ cave and ‘‘Drakos’’ underground river is developed above and below sea-level respectively, in the medium-bedded limestones of the Mani geotectonic unit. The formation and the development of these caves started, most likely, during Middle Pleistocene. Initially, these caves were terrestrial and developed separately. They were connected probably during Holocene through a fi ssure. The development of this united karstic system is controlled by tectonics. ‘‘Selinitsa’’ cave is older than ‘‘Drakos’’. The sequential base levels of karstifi cation demonstrate the continuous sea-level changes during Pleistocene and Holocene, induced by the relative tectonic activity. This united karstic system is characterized by ‘incomplete linkage’ to the sea.