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Enviroscan Ukrainian Institute of Speleology and Karstology

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Speleology in Kazakhstan

Shakalov on 04 Jul, 2018
Hello everyone!   I pleased to invite you to the official site of Central Asian Karstic-Speleological commission ("Kaspeko")   There, we regularly publish reports about our expeditions, articles and reports on speleotopics, lecture course for instructors, photos etc. ...

New publications on hypogene speleogenesis

Klimchouk on 26 Mar, 2012
Dear Colleagues, This is to draw your attention to several recent publications added to KarstBase, relevant to hypogenic karst/speleogenesis: Corrosion of limestone tablets in sulfidic ground-water: measurements and speleogenetic implications Galdenzi,

The deepest terrestrial animal

Klimchouk on 23 Feb, 2012
A recent publication of Spanish researchers describes the biology of Krubera Cave, including the deepest terrestrial animal ever found: Jordana, Rafael; Baquero, Enrique; Reboleira, Sofía and Sendra, Alberto. ...

Caves - landscapes without light

akop on 05 Feb, 2012
Exhibition dedicated to caves is taking place in the Vienna Natural History Museum   The exhibition at the Natural History Museum presents the surprising variety of caves and cave formations such as stalactites and various crystals. ...

Did you know?

That field test is a test run in the field under normal field conditions [16].?

Checkout all 2699 terms in the KarstBase Glossary of Karst and Cave Terms

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What is Karstbase?



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KarstBase a bibliography database in karst and cave science.

Featured articles from Cave & Karst Science Journals
Chemistry and Karst, White, William B.
See all featured articles
Featured articles from other Geoscience Journals
Karst environment, Culver D.C.
Mushroom Speleothems: Stromatolites That Formed in the Absence of Phototrophs, Bontognali, Tomaso R.R.; D’Angeli Ilenia M.; Tisato, Nicola; Vasconcelos, Crisogono; Bernasconi, Stefano M.; Gonzales, Esteban R. G.; De Waele, Jo
Calculating flux to predict future cave radon concentrations, Rowberry, Matt; Marti, Xavi; Frontera, Carlos; Van De Wiel, Marco; Briestensky, Milos
Microbial mediation of complex subterranean mineral structures, Tirato, Nicola; Torriano, Stefano F.F;, Monteux, Sylvain; Sauro, Francesco; De Waele, Jo; Lavagna, Maria Luisa; D’Angeli, Ilenia Maria; Chailloux, Daniel; Renda, Michel; Eglinton, Timothy I.; Bontognali, Tomaso Renzo Rezio
Evidence of a plate-wide tectonic pressure pulse provided by extensometric monitoring in the Balkan Mountains (Bulgaria), Briestensky, Milos; Rowberry, Matt; Stemberk, Josef; Stefanov, Petar; Vozar, Jozef; Sebela, Stanka; Petro, Lubomir; Bella, Pavel; Gaal, Ludovit; Ormukov, Cholponbek;
See all featured articles from other geoscience journals

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Your search for diversity (Keyword) returned 157 results for the whole karstbase:
Showing 1 to 15 of 157
Fungal communities on speleothem surfaces in Kartchner Caverns, Arizona, USA, ,
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Vaughan Michael J. , Maier Raina M. , Pryor Barry M.

Kartchner Caverns, located near Benson, Arizona, USA, is an active carbonate cave that serves as the major attraction for Kartchner Caverns State Park. Low-impact development and maintenance have preserved prediscovery macroscopic cavern features and minimized disturbances to biological communities within the cave.. The goal of this study was to examine fungal diversity in Kartchner Caverns on actively-forming speleothem surfaces. Fifteen formations were sampled from five sites across the cave. Richness was assessed using standard culture-based fungal isolation techniques. A culture-independent analysis using denaturing gradient gel electrophoresis (DGGE) was used to assay evidence of community homogeneity across the cave through the separation of 18S rDNA amplicons from speleothem community DNA. The culturing effort recovered 53 distinct morphological taxonomic units (MTUs), corresponding to 43 genetic taxonomic units (GTUs) that represented 21 genera. From the observed MTU accumulation curve and the projected total MTU richness curve, it is estimated that 51 percent of the actual MTU richness was recovered. The most commonly isolated fungi belonged to the genera Penicillium, Paecilomyces, Phialophora, and Aspergillus. This culturebased analysis did not reveal significant differences in fungal richness or number of fungi recovered across sites. Cluster analysis using DGGE band profiles did not reveal distinctive groupings of speleothems by sample site. However, canonical correspondence analysis (CCA) analysis of culture-independent DGGE profiles showed a significant effect of sampling site and formation type on fungal community structure. Taken together, these results reveal that diverse fungal communities exist on speleothem surfaces in Kartchner Caverns, and that these communities are not uniformly distributed spatially. Analysis of sample saturation indicated that more sampling depth is required to uncover the full scale of mycological richness across spelothem surfaces.

Diversity and dynamics of microarthropods from different biotopes of Las Sardinas cave (Mexico) , ,
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Jos Palaciosvargas, Gabriela Castaomeneses, Daniel A. Estrada

An ecological study of the microarthropod communities from Las Sardinas cave was undertaken. Four different biotopes were studied over the course of a year: bat guano, litter, soil under the chemoautotrophic bacteria colonies and as a control, plain soil without litter or guano. A total of 27,913 specimens of a total of 169 species were collected. Analysis of Variance (ANOVA) showed that there is a significant effect of biotope on the recorded density, and the post hoc Tukey’s test showed that guano is the most different biotope with the highest value of density recorded. The interaction between season and biotope variables was not significant. In the most extreme case, 99 percent of the microarthropods in soil under chemoautotrophic bacteria were mites, mainly in the family Histiostomidae.

Sulfate reducing bacteria in microbial mats: Changing paradigms, new discoveries, 0000,
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Baumgartner Lk, Reid Rp, Dupraz C, Decho Aw, Buckley Dh, Spear Jr, Przekop Km, Visscher Pt,
Sulfate reducing bacteria (SRB) have existed throughout much of Earth's history and remain major contributors to carbon cycling in modern systems. Despite their importance, misconceptions about SRB are prevalent. In particular, SRB are commonly thought to lack oxygen tolerance and to exist only in anoxic environments. Through the last two decades, researchers have discovered that SRB can, in fact, tolerate and even respire oxygen. Investigations of microbial mat systems have demonstrated that SRB are both abundant and active in the oxic zones of mats. Additionally, SRB have been found to be highly active in the lithified zones of microbial mats, suggesting a connection between sulfate reduction and mat lithification. In the present paper, we review recent research on SRB distribution and present new preliminary findings on both the diversity and distribution of [delta]-proteobacterial SRB in lithifying and non-lithifying microbial mat systems. These preliminary findings indicate the unexplored diversity of SRB in a microbial mat system and demonstrate the close microspatial association of SRB and cyanobacteria in the oxic zone of the mat. Possible mechanisms and further studies to elucidate mechanisms for carbonate precipitation via sulfate reduction are also discussed

Quaternary Paleoclimatology of the Black Sea basin, 1979,
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Schrader Hans Joachim,
The occurrence of polyhaline, mesohaline and oligohaline diatom, silicoflagellate, ebridian and chrysomonad populations in late Quaternary Black Sea sediments (DSDP Leg 42B) forms the basis for reconstruction of surface water paleosalinities in the Black Sea basin over the last 3 million years. Four major periods with increased salinites are separated by extended freshwater periods. Based on paleosalinites, indicators of trophic freshwater conditions and changes in diatom species diversity, a correlation is made to the northern Europian glacial--interglacial stratigraphy and this correlation is used to place paleoenvironmental events into a chronostratigraphy. The `synchronous' late Quaternary occurrence of sediments rich in organic carbon in both the Eastern Mediterranean and the Black Sea supports this interpretation.Three different stages in the interaction between the Black Sea and the Eastern Mediterranean Sea are defined: Stage A (exchange of freshwater and marine water similar to the present day flux) during the Holocene, Eemian, Holsteinian and Pliocene; Stage B (freshwater conditions with only occasional marine spills) during the Saalian, the Waalian, the Tiglian and the Praetiglian; and Stage C (freshwater conditions with no inflow of marine waters) during the Weichselian, the Elsterian and Eburonian

Distribution and Habitat Diversity of Subterranean Amphipods in the Rocky Mountains of Colorado, U.S.A., 1981,
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Holsinger John R. , Ward James W.
Subterranean amphipods have been collected from 35 locations on the eastern and western slopes of the Continental Divide in Colorado. All belong to the exclusively subterranean genus Stygobromus. Five species have been identified, two of which are undescribed. Specimens have been collected from (a) the hyporheic zone of rivers, (b) interrupted streams, (c) springs, and (d) seeps at elevations from 1597-2134 m a.s.l. Stygobromus occurs in several habitat types in interrupted drainage basins including sources, seeps, and isolated pools containing leaf detritus. All habitats contained waters which were cool to cold with dissolved oxygen values ranging from 4.3 ppm to fully saturated. Most waters exhibited soft or medium hardness, although one spring containing an undescribed species of Stygobromus had very hard waters (203 ppm bound CO2) and was mildly saline (913 mg/l TDS). There is evidence that the subterranean amphipods are phreatobites which, only under special conditions, establish relatively permanent populations in epigean habitats. Although little is known regarding ecology, zoogeography, or even taxonomy of the subterranean fauna of this region, stygobromid amphipods from the Cordilleran of western North America are apparently represented by fewer well differentiated species per unit area than their congeners from the geobiologicably older Appalachian region of eastern North America where numerous species are found in caves.

Phreatische Fauna in Ljubljansko polje (Ljubljana-Ebene, Jugoslavien); ihre oekologische Verteilung und zoogeographische Beziehungen., 1981,
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Sket Boris, Velkovrh Franci
The phreatic basin of Ljubljansko polje (polje = plain, field) recharges its water supply mainly from the Sava river-bed and at a few other locations where connections with karstic subterranean waters might exist and only up to 15% from precipitation. An important zone of infiltration in the river-bed is the bottom and not the bank which is to a large extent watertight due to organic debris (rests of Sphaerotilus e.g.). The main water-body moves about 10 m/day, there are however some local jets with far higher speeds. Yearly amplitudes of water temperatures are high near the river but in the centre of the plain only a couple of centigrades. Oxygen saturation is in the open river-water 100%, dropping to 40-60% just 1 m into the phreatic. True stygopsammal animals are represented here only by a few species and specimen in spite of the fact, that the interstices in the gravel are mostly filled with finer sediments. Remarkable is also the scarcity of Nematodes and the near absence of Acarina (compare with Danielopol 1976). Only a few specimen of the river benthos (Chironomidae, Tipulidae, Leuctra supp., Baets spp.) penetrate the interstitial water (compare with Ruffo 1961, Danielopol 1976) and only Naididae are more frequent there. However, many epigean animals occur in interstitial waters in the periodically flooded gravel-banks; one can explain this with oscillations of the water level. Some epigean animals (creno- and troglophilic) are quite regularly represented in the phreatic near the river, but have not been found in the river-bed. The distribution of phreatic species within the studied water-body seems to be controlled mainly by the presence of food supplies and the consequent competition among species. The same is true for the speed of the water current and some other factors which are less easily defined. The characteristics of the substratum as well as O2-saturation and other characteristics of the water seem to have little influence on the fauna. The energetically (food-) rich neighbourhood of the river is inhabited by a number of species in quite dense populations while the central parts of the phreatic water body exhibit a great poverty of species and of specimen. However, some species live here, which don't occur in the presence of larger food supplies and of greater competition (Niphargus serbicus). The higher current speed seems to prevent settlement of some species (Cyclopoida, Proasellus deminutus) while some are bound to such habitats (Proasellus vulgaris). Some species exhibit a high degree of euryvalency inside the stygopsephale habitats (Niphargus longidactylus e.g.), while some are highly specialized. Some of them form dense populations (comparatively dense even in energetically poor places) while others exhibit even in most favourable conditions very low densities (Niphargus jovanovici multipennatus). The present fauna is zoogeographically very diverse. Some species are distributed throughout Europe; some reach from Central Europe to the borders of Dinaride Karst (Bogidiella albertimagni) and some even penetrate it (Trichodrilus pragensis, Acanthocyclops kiefer). Bogidiella semidenticulata. Niphargus pectinicauda, Hadziella deminuta seem to be limited to the higher reaches of the Sava River. All of the above mentioned animals live regularly in interstitial waters and only sporadically in karstic hypogean waters. Niphargus stygius is here the only animal of a certainly karstic provenience; inside the plain it is limited to a completely special habitat. It is very likely that the entire Proasellus-deminutusgroup has developed in interstitial waters of larger plains which are in contact with karstic areas; some species penetrated from the plains into the karst rather than the reverse. To the contrary (judging from the distribution of the genera) karstic waters seem to be the cradle of Hauffenia and Hadziella. Such a sharp delimitation between cave- and interstitial fauna resp. in this area is very noteworthy. Both faunas live here in abundance and in close contact. It is very probable that particularly high competition and specialization of both faunas, caused by their richness and diversity, prevent mixing of species.

Le Vercors : un massif de la moyenne montagne alpine, 1984,
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Delannoy, J. J.
THE VERCORS: A MASSIF OF MIDDLE ALPINE MOUNTAIN - The Vercors is a forested massif of middle mountain, in the French Northern Prealps. The characteristic of the Vercors massif is the thick and massif urgonian limestones, which underlines a folded structure. The Vercors quickly hold the attention of speleologists, who had put forward the density and diversity of karstic aspects since the beginning of the century. Landforms display glacial karstic landscapes more or less damaged in terms of bio-climatic levels of mounts and depressions landscapes and of deep water gaps, which main flows of the massif are flowing in. The notion of the morphoclimatic heritage applies to the underground karst for which glacial quaternary episodes have been determinant in the large systems genesis (gouffre Berger - Scialet de la Fromagre, Antre des Damns, Combe de Fer...). The study of underground deposits allows to bring up-to-date various period of karst development; a preglacial stage (e.g. upper levels of Cuves de Sassenage, Gournier, Coufin-Chevaline...). The study of the current dynamic shows that the Vercors as an important karstic ablation, between 120 to 170mm/ky. The karstic dynamic exerts mainly on the superficial slab of the massif (from 80 to 50% of the whole ablation). The Vercors can be considered as the best example of calcareous massif in temperate middle mountains, thanks to the combination of various favourable parameters: pure karst rocks, morpho-climatic episodes not constraining, and a high karstic dynamic.

Un haut plateau jurassien : le plateau des Molunes (Jura), 1984,
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Neyroud, M.
LARGE KARSTIC DEPRESSIONS AND ANTICLINAL STRUCTURES ON A HIGH JURASSIC TABLE-LAND: THE SOUTH OF THE PAYS MOLUNOIS (JURA, FRANCE) - On a high Jurassic table-land, the Pays molunois, close to the Monts Jura (Crt de la Neige, 1717m), large karstic depressions are studied in connection with anticline structures. Along a single anticline crest, a great diversity of forms is explained by the character of strike-slip faults (direct tectonic control), the glacial heritage (regional ice-cap), the tardiglacial evolution (peat-bogs or molunes), the lithological context (minor morphology) and at last unequal anthropic action. The morphogenesis of an original type of small anticlinal valleys, the closed anticline valleys-poljes is considered dating from geomorphologic evolution of the area and particular tectonic context.

Premires observations morphologiques et splologiques sur le karst haut-alpin du Tennengebirge (Salzburg, Autriche), 1987,
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Audra, Ph.
THE MASSIF OF TENNENGEBIRGE (SALZBURG, AUSTRIA) - The Tennengebirge, in the Salzburg limestone Alps, are characterised by an important high alpine karst, containing some of the deepest austrian caves (Batman Hhle: -1219m, Schneeloch: 1086m, Platteneck: -937m, Gipfel Loch: -852m). The phreatic zone locates at 720m asl. The main springs are situated on the northern side of the mountain. A description of the Vordere Pitschenberg Valley has been sketched illustrating a large diversity and underground, of actual or inherited character.

Le karst jurassique du rebord subalpin dans les Alpes-Maritimes, 1989,
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Mangan, C.
Jurassic karst of the subalpine area in Maritime Alps Department (France) - The jurassic carbonated sediments represent the principal karst aquifer in front of the alpine range. This area is subdivided in three different sectors by the tectonic and paleogeographic evolution: subalpine arc of Castellane (Grasse Prealps), subalpine arc of Nice, Provence foreland. Through the comparative study of these areas, it appears a real diversity about the karst evolution, the geometry about the karst evolution, the geometry of waters reservoirs, the distribution of the underground flows, and therefore about the water supply and exploitation.

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Ray J. A. , Odell P. W. ,
This study outlines an improved method, DIVERSITY, for delineating and rating groundwater sensitivity. It is an acronym for Dlspersion/VElocity-Rated SensitivITY, which is based on an assessment of three aquifer characteristics: recharge potential, flow velocity, and flow directions. The primary objective of this method is to produce sensitivity maps at the county or state scale that illustrate intrinsic potential for contamination of the uppermost aquifer. Such maps can be used for recognition of aquifer sensitivity and for protection of groundwater quality. We suggest that overriding factors that strongly affect one or more of the three basic aquifer characteristics may systematically elevate or lower the sensitivity rating. The basic method employs a three-step procedure: (1) Hydrogeologic settings are delineated on the basis of geology and groundwater recharge/discharge position within a terrane. (2) A sensitivity envelope or model for each setting is outlined on a three-component rating graph. (3) Sensitivity ratings derived from the envelope are extrapolated to hydrogeologic setting polygons utilizing overriding and key factors, when appropriate. The three-component sensitivity rating graph employs two logarithmic scales and a relative area scale on which measured and estimated values may be plotted. The flow velocity scale ranging from 0.01 to more than 10,000 m/d is the keystone of the rating graph. Whenever possible, actual time-of-travel values are plotted on the velocity scale to bracket the position of a sensitivity envelope. The DIVERSITY method was developed and tested for statewide use in Kentucky, but we believe it is also practical and applicable for use in almost any other area

The last glacial/interglacial record of rodent remains from the Gigny karst sequence in the French Jura used for palaeoclimatic and palaeoecological reconstructions, 1995,
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Chaline J, Brunetlecomte P, Campy M,
A multidisciplinary approach has produced an exceptional chronological log of climatic patterns for the Upper Pleistocene sequence of Gigny Cave (Jura, France) covering the Pre-Eemian, Eemian Interglacial, Middle Glacial and Upper Pleniglacial, as well as a part of the Holocene. Multivariate analysis (correspondence and component analysis) of rodent associations from the sequence is used here to characterize the different climatic stages in terms of relative temperature, plant cover and moisture. Faunal analysis establishes: (1) positive and negative correlations among the variations of the different species; (2) the significance of axis 1 (component analysis) which, in terms of temperature, opposes cold environments with contrasted continental biotopes; (3a) the significance of axis 2 (component analysis), which reflects vegetation patterns ranging from open to closed habitats; (3b) the significance of axis 3 (component analysis), which expresses trends in moisture; (4) various correlations between faunal and climatic parameters (temperature, plant cover and moisture); (5) evaluation of faunal diversity (Shannon index ranging from 0.74 to 2.27) showing that diversity increases with temperature and the complexity of vegetation, but is not sensitive to moisture. Lastly, the comparison of multivariate methods with the weighted semi-quantitative Hokr method shows the complementarity of the two approaches, the first methods quantifying climatic parameters while the second seems to provide more precise evaluations of the main seasons of rainfall

Searching for extinction/recovery gradients: the Frasnian-Famennian interval, Mokra Section, Moravia, central Europe, 1996,
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Cejchan P, Hladil J,
A series of ancient seafloors colonized by diverse organisms has been documented from the Upper Devonian rocks of the Western Mokra Quarry. Situated in the southern tectonic closure of the Moravian Karst, the Frasnian-Famennian shallow carbonate ramps exhibit both Rhenish and Ukrainian affinities. Reconstruction of palaeo-sea floor horizons results in a series of 28 quadrats sufficient for further evaluation. Eighty-five taxa involved were scrutinized for abundance, occupied area, skeletal mass production and biomass production. The aim of the study was to determine whether the observed sequence of quadrats can be distinguished from a random one, and to discover any possible unidimensional gradient as a latent control. Monte Carlo simulations and a graph theoretical approach were utilized. Although the raw data seemed chaotic, the simulations demonstrated the observed sequence is not random. A significant influence of a hidden control is thus suggested. Fifteen characteristics of quadrats (e.g. diversity, number of taxa, vertical stratification of community, number of patches) were utilized for final interpretation. The gradient reconstructed by TSP algorithm reveals a significant crisis within the uppermost part of the Amphipora-bearing limestone

Laquifre de la source du Lez : un rservoir deau et de biodiversit, 1997,
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Malard Florian, Gibert Janine, Laurent Roger
The Lez spring is the main source of drinking water for the inhabitants of the city of Montpellier. This spring has been exploited since the eighteenth century but the amount of groundwater pumped has markedly increased over the last 30 years. This karst harbours an extremely diversified community of groundwater species (at least 37 species) that is a several Million-year-old heritage. Overpumping induces a loss of habitats by lowering the water table during periods of low groundwater recharge. It also results in an artificial fragmentation of mesohabitats by increasing the hydraulic disconnection of different regions in the saturated zone. Thus, overpumping may strongly affect the groundwater fauna but few data are available yet to evaluate the potential loss of biodiversity. There is clearly a need to integrate studies of groundwater fauna within the framework of interdisciplinary groundwater monitoring, management and/or protection programmes.

Long term stability of a terrestrial cave community, 1997,
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Carchini Gianmaria, Di Russo Claudio, Luccarelli Marco, Rampini Mauro, Sbordoni Valerio
We report data on the spatial structure and seasonal variation of the community of Valmarino cave, a medium sized sandstone cave, located a few kilometres from the coast line, in Central Italy. Due to both its habitat features and its relatively recent geological history, Valmarino cave is only inhabited by terrestrial, troglophilic elements, i.e facultative cave dwellers. By means of monthly censuses and density plot estimates we have investigated species abundance, diversity and their spatial organization, by considering separately samples from different cave sectors. Homogeneous sampling design allowed to compare series of samplings performed in 1974 and 1994. On the whole 21 arthropods and one snail species constitute the cave community. Ordination plots resulting from correspondence analyses of monthly samples outline a distinct spatial and temporal structure. Two main sub-communities can be identified: a inner subcommunity, mainly represented by eu-troglophilic species, showing a remarkable stability throughout the year and an outer sub-community, mainly represented by sub-troglophilic species, showing strong seasonal variation. Both spatial and temporal vectors show similar importance in shaping the community structure. An interesting result of this study is the long term stability of both spatial and seasonal components of the community structure which remained almost identical after 20 years, as shown by the comparison of ordination plots obtained from 1974 and 1994 sampling series. Therefore this study provides empirical evidence of a frequently hypothesised, albeit never demonstrated feature of the cave ecosystem.

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