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Speleology in Kazakhstan

Shakalov on 04 Jul, 2018
Hello everyone!   I pleased to invite you to the official site of Central Asian Karstic-Speleological commission ("Kaspeko")   There, we regularly publish reports about our expeditions, articles and reports on speleotopics, lecture course for instructors, photos etc. ...

New publications on hypogene speleogenesis

Klimchouk on 26 Mar, 2012
Dear Colleagues, This is to draw your attention to several recent publications added to KarstBase, relevant to hypogenic karst/speleogenesis: Corrosion of limestone tablets in sulfidic ground-water: measurements and speleogenetic implications Galdenzi,

The deepest terrestrial animal

Klimchouk on 23 Feb, 2012
A recent publication of Spanish researchers describes the biology of Krubera Cave, including the deepest terrestrial animal ever found: Jordana, Rafael; Baquero, Enrique; Reboleira, Sofía and Sendra, Alberto. ...

Caves - landscapes without light

akop on 05 Feb, 2012
Exhibition dedicated to caves is taking place in the Vienna Natural History Museum   The exhibition at the Natural History Museum presents the surprising variety of caves and cave formations such as stalactites and various crystals. ...

Did you know?

That oxbow is abandoned loop of a stream course, original usage, applied to surface rivers, describes short-circuited meander loops but in caves the term is applied to dry loop passages of any shape and origin [9].?

Checkout all 2699 terms in the KarstBase Glossary of Karst and Cave Terms

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Featured articles from Cave & Karst Science Journals
Chemistry and Karst, White, William B.
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Featured articles from other Geoscience Journals
Karst environment, Culver D.C.
Mushroom Speleothems: Stromatolites That Formed in the Absence of Phototrophs, Bontognali, Tomaso R.R.; D’Angeli Ilenia M.; Tisato, Nicola; Vasconcelos, Crisogono; Bernasconi, Stefano M.; Gonzales, Esteban R. G.; De Waele, Jo
Calculating flux to predict future cave radon concentrations, Rowberry, Matt; Marti, Xavi; Frontera, Carlos; Van De Wiel, Marco; Briestensky, Milos
Microbial mediation of complex subterranean mineral structures, Tirato, Nicola; Torriano, Stefano F.F;, Monteux, Sylvain; Sauro, Francesco; De Waele, Jo; Lavagna, Maria Luisa; D’Angeli, Ilenia Maria; Chailloux, Daniel; Renda, Michel; Eglinton, Timothy I.; Bontognali, Tomaso Renzo Rezio
Evidence of a plate-wide tectonic pressure pulse provided by extensometric monitoring in the Balkan Mountains (Bulgaria), Briestensky, Milos; Rowberry, Matt; Stemberk, Josef; Stefanov, Petar; Vozar, Jozef; Sebela, Stanka; Petro, Lubomir; Bella, Pavel; Gaal, Ludovit; Ormukov, Cholponbek;
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Your search for rodents (Keyword) returned 15 results for the whole karstbase:
Showing 1 to 15 of 15
On the knowledge of Mammal fauna of the Banat Caves (Romania)., 1967, Botosaneanu Lazare, Negrea Alexandrina, Negrea Stefan
The authors assembled from about 70 caves a rich collection of osteological material and specimens of living or fossil mammals. A list of the caves is given with an enumeration of the identiied species for each cave. Under each species the caves which supplied the material are listed. This is followed by an inventory of the osteological material and by observations on the living animals (especially bats). Fifty-three mammal species (fossil and living) were accurately determined (14 carnivores, 6 artiodactyls, 1 lagomorph, 10 rodents, 3 insectivores, and 19 bats).

Late Pleistocene microtine rodents from Snake Creek Burial Cave, White Pine County, Nevada, 1998, Bell Cj, Mead Ji,
A total of 395 microtine rodent specimens recovered from Snake Creek Burial Cave (SCBC) are referred to Microtus SP. and Lemmiscus curtatus. Radiocarbon and Uranium series dates indicate an ae for these fossils of between 9460 160) yr. B.P. and 15,1000 700 yr, B.P. The sample of lower first molars of Lemmiscus includes 4-, 5-, and B-closed triangle morphotypes. Earlier reports of the 4-closed triangle morphotype are from Irvingtonian deposits in Colorado, Nevada, and New Mexico and from early Rancholabrean deposits in Washington. The morphotype is not known in living populations of Lemmiscus. SCBC specimens constitute the youngest record of the 4-closed triangle morphotype and are the only-specimens reported item the late Rancholabrean. Thc time of disappearance of Lemmiscus with this molar morphology is unknown, but populations with this morphotype possibly became extinct at or near the end of the Pleistocene

A succession of Miocene rodent assemblages from fissure fillings in southern France: palaeoenvironmental interpretation and comparison with Spain, 1999, Aguilar Jp, Escarguel G, Michaux J,
An Early to Late Miocene sequence of rodent assemblages from southern France has been quantitatively studied. The resulting pattern seems very similar to a contemporary sequence from central Spain (Calatayud-Teruel Basin). The fossil mammal-bearing localities are of different types: mainly karst infills in France and localities situated in sedimentary basins in Spain. In order to interpret the fossil record, a comparison has been made between southern France faunas of similar age but collected in karst infills and in basin deposits. There seems to be no difference between the two kinds of faunas and thus there is no indication that karst infills systematically give a picture of drier and more open environments. Both types of localities may give a similar relative abundance of taxa and when differences exist they can be attributed to local conditions. The comparison between southern France and the Calatayud-Teruel Basin (central Spain) shows that: (1) similar trends occurred in the two areas; (2) differences between spectra were more important during the late Early Miocene than during the Middle Miocene; (3) the shift between the late Early Miocene and the Middle Miocene environments in southern France does not seem to be correlated with. a general drop in temperatures as inferred from the analysis of central Spain faunas. (C) 1999 Elsevier Science B.V. All rights reserved

The small vertebrate fauna (Rodents, Insectivores, and Reptiles) of Šandalja 1A (Istria, Croatia), 2001, Aguilar Jeanpierre, Crochet Jeanyves, Michaux Jacques, Mihevc Andrej, Paunovič, Maja

Is described under the name of Šandalja 1A, a Lower to Middle Pleistocene fauna of small vertebrates including rodents, insectivores and reptiles. Extracted from a bone breccia found in 1999 in the Šandalja quarry near Pula, its accurate localization with respect to the previously known bone breccia of Šandalja 1 is not known. Nevertheless this dating - a Biharian age - is congruent with the younger age now advocated for the fauna of large mammals of Šandalja 1 and its associated chopper.

A possible heptaxodontine and other caviidan rodents from the quaternary of Jamaica, 2003, Macphee R. D. E. , Flemming C. ,
New World hystricognath rodents (parvorder Caviida) easily qualify as the most diversified members of the nonvolant Quaternary land mammal fauna of the West Indies. This paper describes three intriguing but problematic representatives of this group from Jamaican cave deposits. The first is the holotype (and still the only) specimen of Alterodon major from Wallingford Roadside Cave, a taxon that continues to generate controversy because specialists disagree as to its placement within Caviida. We reject the argument that it should be placed in Octodontidae and reaffirm the high probability that it is a clidomyine. The second fossil is a large proximal femur, apparently recovered from Sheep Pen locality near Windsor (Trelawney Parish) in the 1960s. Much larger than the femur of Clidomys (previously thought to be Jamaica's largest Quaternary mammal), in size and morphology the new fossil somewhat resembles femora of the eastern Caribbean heptaxodontine Amblyrhiza. Although firm allocation is not possible, the Sheep Pen femur is possibly that of a megafaunal caviidan. The third fossil described in this paper is the jaw of a previously unknown caviidan from a dated end-Pleistocene cave context in Portland Ridge (Jackson's Bay, Clarendon Parish). Xaymaca fulvopulvis, new genus and species, differs from all West Indian caviidan species presently known. The jaw is well preserved but retains only the incisor and premolar (the latter in a very worn state). The few features for which the new species can be usefully analyzed and compared to caviidan groups represented in the West Indian Cenozoic (capromyids, heteropsornyines, heptaxodontines, and clidomyines) are largely indecisive from a systematic perspective. However, on balance the strongest indicators seem to lie with the 'giant' heptaxodontines of the central and eastern Caribbean (the grouping composed of Amblyrhiza, Elasmodontomys, and possibly Quemisia), and despite its diminutive size Xaymaca is tentatively placed within that group. It is increasingly apparent that much still remains to be learned about the origin and history of the land mammal fauna of Jamaica

Etude des palokarsts des environs de Saint-Remze (Ardche, France) : mise en vidence dune rivire souterraine fossilise durant la crise de salinit messinienne, 2005, Martini, Jacques
Paleokarst investigation near Saint-Remze, Ardche, France discovery of an underground river fossilized during the Messinian salinity crisis - The paleokarst features studied in this paper are hosted in Lower Cretaceous limestone and generally appear as filled caves, subsequently de-roofed by denudation. The most important of them forms a sequence of segments developed at a relatively constant elevation of 360-380 m above sea level and can be traced over a length of 5.2 km. The ancient cave passages generally appear as soil covered bands, 5 to 20 m in width and limited on both sides by limestone outcrops. At surface the nature of the cave filling is revealed mainly by scattered blocks: calcite from speleothems and calcified clay, silt, sand and breccia. In the best preserved places, the earth band lies in a trench, where the walls may display a cave morphology and where the filling is often exposed in a undisturbed state. Three types of detrital cave filling have been identified, which in stratigraphical order are as follows: 1) Beige-grey silt, sand and microconglomerate of immature alluvials, with elements of Paleozoic granites and metamorphic rocks, and Upper Miocene volcanics, both originating from the Cvennes Mountains 30 km to the NW. The lithological composition is comparable to the recent alluvials of the Paleokarst investigation near Saint-Remze, Ardche, France discovery of an underground river fossilized during the Messinian salinity crisisArdche River, which is flowing a few kilometers to the SW and is deeply entrenched into a canyon at elevations of 40 to 80 m. The karst context, combined with the biostratigraphical data obtained from rodent molars in the alluvials, suggests an Uppermost Miocene age, comprised between ~5.8 and ~5.45 Ma. 2) Red mature alluvials and colluvials originating from local reworking of surficial karst residuals. At one spot they gave a paleontological age of 3.6 to 3.0 Ma, but from the local karst context one may expect ages from final Miocene to Pleistocene in other spots. 3) Monogenic breccia generated from wall gelifraction, which is Pleistocene after rodent molars found in two places. The paleocave is visualised as formed by an underground stream fed from swallow-holes on the bank of the Ardche River, when it was flowing more than 300 m higher than its actual bed. With regard to its relatively constant elevation and a discordant relationship with the country rock bedding, it is interpreted as a vadose cave controlled by a paleo-water-table. The other fillings (2 and 3) were deposited during subsequent vadose speleogenesis and after considerable water-table lowering. The elevation of this fossilised underground river coincides fairly well with the pre-salinity crisis abandonment surface (5.52 Ma), which is evidenced in the area by high perched gravel relics. The end of the speleogenesis could have taken place just before this event (~5.6 Ma) or at an age not younger than ~5.45 Ma. In the latter possibility, speleogenesis had to be working before the regressive erosion generated by the drastic lowering of the Mediterranean Sea [5.52 to 5.33 Ma, Clauzon et al, 2005 ] reached the area and de-watered the deep karst aquifer. This fossil underground river provides also information about the morphological evolution of the area. For instance the nature of the immature alluvials suggests that the torrential regime of the Ardche was about the same than today. It also indicates that the important and famous cave systems in the area (Grotte de Saint-Marcel, Aven dOrgnac, Systme de Foussoubie, Grotte Chauvet), which are developed at lower elevations, cannot be older than ~5.6 Ma and most likely formed mainly during the Plio-Pleistocene, although most of them have been initiated during the salinity crisis.

New palaeontological assemblage, sedimentological and chronological data from the Pleistocene Ma U'Oi cave (northern Vietnam), 2006, Bacon Am, Demeter F, Rousse S, Long Vt, Duringer P, Antoine Po, Thuy Nk, Mai Bt, Huong Ntm, Dodo Y,
This paper describes recent material gathered during the second fieldwork at Ma U'Oi in November 2002 by a Vietnamese-French-Japanese team. The Ma U'Oi cave, located in the province of Hoa Binh (60 km SW from Hanoi), northern Vietnam, belongs to a karstic network developed in Triassic dark-grey limestones.The cave is filled with coarse-grained breccias containing numerous fossil remains, partially preserved at several loci inside the cave (wall, vault and ground). We describe new teeth which confirm the occurrence of mammal taxa already mentioned at Ma U'Oi (Bacon et al., 2004)[Bacon, A-M., Demeter, F., Schuster, M., Long, V.T., Thuy, N.K., Antoine, P-O., Sen, S., Nga, H.H., Huong, N.T.M., 2004. The Pleistocene Ma U'Oi cave, northern Vietnam: palaeontology, sedimentology and palaeoenvironments. Geobios 37, 305-314], while others, mainly microvertebrates, emphasize the occurrence of new species for the Pleistocene of Vietnam. We report here, for the first time, the occurrence of these microvertebrates of different groups (primates, rodents, insectivores, small reptiles and amphibians) in the faunal assemblage. Among mammal taxa, the presence of one more hominid affiliated to archaic Homo is also attested by our findings. U/Th dating carried out on 2 samples extracted from breccia speleothems confirms the biochronological estimate, with fossiliferous fillings ranging from late Middle Pleistocene to Late Pleistocene


For the first time in the Classical Karst, paleontological data enabled to match the magnetostratigraphic record precisely with the geomagnetic polarity timescale in two studied sites: (i) a series of speleothems alternating with red clays in Račiška pečina Cave (Matarsko podolje), and (ii) an unroofed paleocave of the Črnotiče II site (Podgorski kras Plateau) completely filled by fluvial clastic sediments covered by speleothems. The later sites are also characterized by a rich appearance of fossil tubes of autochthonous stygobiont serpulid Marifugia cavatica. The vertebrate record is composed mostly of enamel fragments of rodents and soricomorphs. Absence of rootless arvicolids as well as taxonomic composition of the mammalian fauna suggests the Pliocene age of both sites. For (i) Račiška pečina (with Apodemus, cf. Borsodia) it was estimated to middle to late MN17 (ca 1.8–2.4 Ma), while (ii) the assemblage from Črnotiče II (with Deinsdorfia sp., Beremedia fissidens, Apodemus cf. atavus, Rhagapodemus cf. frequens, Glirulus sp., Cseria sp.) is obviously quite older: MN15–MN16 (ca 3.0–4.1 Ma). In respect to congruence of biostratigraphic and paleomagnetic data and a reliable sedimentary setting of the samples we propose to apply the respective datum also as the time of one ancient speleogenetic phase in the Classical Karst.

Early late Pliocene paleokarstic fillings predating the major Plio-Pleistocene erosion of the Quercy table, SW-France, 2007, Aguilar J. P. , Michaux J. , Pé, Lissié, T. , Sigé, B.

This evolution consists, first in the elaboration of the underground net systems, then in a long polyphased process of filling – emptying the karstic voids, according to the up and down base level changes that occurred almost continuously during the first half of the Tertiary era. The filling sediments are mostly vadose clay deposits, the various ages of which being established from the study of their fossil vertebrate contents (for latest accounts see Pélissié & Sigé 2006). Then, since latest Oligocene times, the Quercy platform was covered with prograding lacustrine sediments of the Aquitaine Basin. Finally, the whole structure was strongly worn down by the so- called Plio-Pleistocene erosional phase: the previously deep underground system became closer to the surface, and was exposed both to erosion and widening, but also Plio-Pleistocene fillings occurred as shown by the fossils they include (Crochet et al. 2006). Among the latter are rare Late Pliocene and Plio-Pleistocene tooth specimens.

Impacts of Alterations of Organic Inputs on the Bacterial Community within the sediments of Wind Cave, South Dakota, USA., 2009, Chelius M. K. , Beresford G. , Horton H. , Quirk M. , Selby G. , Simpson R. T. , Horrocks R. , Moore J. C.
Wind Cave (WICA) in the Black Hills of South Dakota, like many mostly dry caves in temperate regions is an energy-starved system. The biotic communities that reside in these systems are low in diversity and simple in structure, and sensitive to changes in external inputs of organic matter. Caves open to tourist traffic offer an opportunity to study the impacts of organic matter amendments in the form of human and rodent hair and dander, clothing lint, material from rodent activity (nesting materials and feces), and algal growth in and around artificial lighting. This study reports on the impacts of carbon amendments from humans and rodents on the bacterial and archaeal communities within the sediments of WICA from annual surveys and from a manipulative study that added lint (L; cellulose plus rodent dander and rodent hair), rodent feces (F), and a combination of both (LF). The survey confirmed that bacterial biomass was higher in regions of the cave with the highest rates of lint (hair and natural clothing fibers) input. The manipulative study found that organic amendments in the forms of lint (L) and rodent feces (F) altered the WICA bacterial community structure in both abundance and diversity, with the combined lint and feces (LF) amendment having the most significant response. The high similarity of the LF and L communities suggests that the cave bacterial community is more carbon than nitrogen limited. The implication of cave development to management practices is immediate and practical. Even small amounts of lint and organic matter foreign to cave bacteria significantly compromise the integrity of the endemic community resulting in the replacement of undescribed species by assemblages with at best, unknown impacts to natural cave features.

Paleokarst investigation near Saint-Remèze, Ardèche, France: discovery of an underground river fossilised during the Messinian salinity crisis, 2011, Martini, J. E. J.

A number of paleokarst fillings have been investigated. The most important of them represents an ancient underground river. It appears as a sequence of filled passage segments, which have been de-roofed by karst denudation. These segments are developed at 360 to 380 m above sea level and have been followed for 5.2 km. Three distinct cave fillings were put into evidence: 1) beige micaceous silts and sands which represent exogenic immature alluvials and were dated with rodent bones as Uppermost Miocene; 2) mature red clay and sandstone of local origin, whose age might vary from Uppermost Miocene to Recent; 3) monogenic breccia generated by wall gelifraction during the Pleistocene. The petrographic composition of the immature alluvials is similar to the one of the Ardèche River which flows in the vicinity, but deeply entrenched in a canyon, at the altitude of 60-70 m ASL. Therefore, the paleo-underground river was fed by ponors located on the bank of the Ardèche River, when it was flowing more than 300 m higher than its present bed. The weak variations in elevation of the fossil channel suggest a development within the immediate vicinity of the water table. The biostratigraphic age of the immature alluvials as well as the paleokarstic context suggest that the cave was still active ~5.6 to 5.45 Ma ago. In this timespan falls the drastic dryout of the Mediterranean Sea and the beginning of the incision of the Messinian canyons in this area. In general, this fossil water-table cave is also informative on the morphological evolution of the Ardèche Karst and underline the usefulness of palaeontology in dating speleogenesis.

Pleistocene edible dormice (Rodentia Mammalia) from Slovenia, and their relations to the present day Glis glis (Linnaeus 1766), 2011, Aguilar Jeanpierre, Michaux Jacques

A Pleistocene new material of dormice (Genus Glis) is described.
Three morphological species are recognized on the basis of size and morphology of the teeth: Glis sackdillingensis Heller, 1930, Glis mihevci nov. sp., and Glis perkoi nov. sp. The two new species, larger than G. sackdillingensis, are morphologically less evolved than the present day Glis glis of Slovenia, which has larger teeth.

A large Cervidae Holocene accumulation in Eastern Brazil: an example of extreme taphonomical control in a cave environment, 2012, Hubbe Alex, Auler Augusto S.

A remarkable cervid bone accumulation occurs at a single passage (named Cervid Passage; CP) at Lapa Nova, a maze cave in eastern Brazil. CP lies away from cave entrances, is a typical pitfall passage and contains bone remains of at least 121 cervids, besides few bats, peccaries and rodents remains. There is no evidence of water (or sediment) flow at the site and in general bones lack post depositional alterations and display anatomical proximity, suggesting that the majority of the remains found inside CP (mainly cervids) are due to animals that after entering the cave got trapped in the site. Observations suggest that two entrances could have provided access to cervids (and the few other animals, besides bats), either by falling inside the cave or by entering by their own free will. Once inside the cave, the maze pattern would make route finding difficult, and of all passage intersections, only the one leading to CP would result in a non-return situation, starving the animal to death. Radiocarbon dates suggest that animal entrapment occurred during at least 5 thousand years, during the Holocene. The reasons why mainly cervids were found are unknown but they are probably related to the biology of this group coupled with the fact that caves provide several specific taphonomic processes that may account for a strong bias in bone accumulation. Indeed, the frequent occurrence of Cervidae in both the fossil and sub-fossil record in Brazilian caves may be related to an overall high faunal abundance or may suggest that these animals were especially prone to enter caves, perhaps in search of nutrients (as cave saltpetre) or water.

Emine-Bair-Khosar Cave in the Crimea, a huge bone accumulation of Late Pleistocene fauna, 2013, Ridush . , Stefaniak K. , Socha P. , Proskurnyak Y. , Marciszak A. , Vremir M. , Nadachowski A.

The Crimean Mountains are well known from the abundance of Middle and Late Palaeolithic sites and palaeontological remains recovered from cultural layers in caves and rockshelters. The fossil-bearing deposits of Emine-Bair-Khosar Cave, located at the elevation of 1000 m on the Chatyrdag Plateau, yielded a very diverse and numerous vertebrate remains that widen the knowledge of Late Pleistocene faunal diversity in the Crimea. The assemblage comprised in total almost 50 species of vertebrates. Studies included geomorphological, geological and stratigraphic analyses as well AMS 14C dating. Faunal remains were present in ten palaeontological sites. The main bone accumulation (section Ba2) was deposited during Middle Valdai or Vytachiv (MIS 3) interstadial, and including a long time gap, to the end of the Pleistocene and the Holocene. Comparison of the Emine-Bair-Khosar fauna with vertebrate faunas of other Crimean sites showed a remarkable stability in the faunal composition and frequency during the whole MIS 3 interstadial. Steppe and other open-country species dominated in the compared assemblages, while boreal-tundra species were far less numerous. Inhabitants of forests, including red deer and some rodents, were stable members of fossil assemblages.

Late Quaternary Caviomorph Rodents (Rodentia: Hystricognathi) from Ceara State, Northern Brazil, 2013, De Oliviera P. V. , Ribeiro A. M. , Kerber L. , Lessa G. , Viana M. S.

In this paper we report the first remains of caviomorph rodents from the karst of the Parque Nacional de Ubajara, Ceara´ State, northeastern Brazil, collected with precise stratigraphic and radiometric control. The material is derived from levels with thermoluminescence dating of about 8,000 years BP, corresponding to the early Holocene. In these levels, we found remains of Kerodon rupestris Wied, 1820, cf. Dasyprocta Illiger, 1811 and Thrichomys Trouessart, 1880. The data here reported contribute to the knowledge of Brazilian Quaternary rodents and show the potential of the studied area for fossils.

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