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Speleology in Kazakhstan

Shakalov on 04 Jul, 2018
Hello everyone!   I pleased to invite you to the official site of Central Asian Karstic-Speleological commission ("Kaspeko")   There, we regularly publish reports about our expeditions, articles and reports on speleotopics, lecture course for instructors, photos etc. ...

Speleology in Kazakhstan

Shakalov on 04 Jul, 2018
Hello everyone!   I pleased to invite you to the official site of Central Asian Karstic-Speleological commission ("Kaspeko")   There, we regularly publish reports about our expeditions, articles and reports on speleotopics, lecture course for instructors, photos etc. ...

Speleology in Kazakhstan

Shakalov on 11 Jul, 2012
Hello everyone!   I pleased to invite you to the official site of Central Asian Karstic-Speleological commission ("Kaspeko")   There, we regularly publish reports about our expeditions, articles and reports on speleotopics, lecture course for instructors, photos etc. ...

New publications on hypogene speleogenesis

Klimchouk on 26 Mar, 2012
Dear Colleagues, This is to draw your attention to several recent publications added to KarstBase, relevant to hypogenic karst/speleogenesis: Corrosion of limestone tablets in sulfidic ground-water: measurements and speleogenetic implications Galdenzi,

The deepest terrestrial animal

Klimchouk on 23 Feb, 2012
A recent publication of Spanish researchers describes the biology of Krubera Cave, including the deepest terrestrial animal ever found: Jordana, Rafael; Baquero, Enrique; Reboleira, Sofía and Sendra, Alberto. ...

Caves - landscapes without light

akop on 05 Feb, 2012
Exhibition dedicated to caves is taking place in the Vienna Natural History Museum   The exhibition at the Natural History Museum presents the surprising variety of caves and cave formations such as stalactites and various crystals. ...

Did you know?

That carbonic acid dissolution is dissolution of calcium carbonate by carbon dioxide in aqueous solution, loosely termed carbonic acid, is the dominant reaction in karst processes, including speleogenesis. the reaction can be considered in several ways but it is most simply represented as: caco3 + co2 + h2o = ca(hco3)2 the reaction is reversible. the solution containing the dissolved reaction product, usually termed calcium bicarbonate, can lose carbon dioxide to the atmosphere and precipitate calcium carbonate. this process is responsible for the development of speleothems underground and tufa or travertine at the surface [9].?

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KarstBase a bibliography database in karst and cave science.

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Chemistry and Karst, White, William B.
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Geochemical and mineralogical fingerprints to distinguish the exploited ferruginous mineralisations of Grotta della Monaca (Calabria, Italy), Dimuccio, L.A.; Rodrigues, N.; Larocca, F.; Pratas, J.; Amado, A.M.; Batista de Carvalho, L.A.
Karst environment, Culver D.C.
Mushroom Speleothems: Stromatolites That Formed in the Absence of Phototrophs, Bontognali, Tomaso R.R.; D’Angeli Ilenia M.; Tisato, Nicola; Vasconcelos, Crisogono; Bernasconi, Stefano M.; Gonzales, Esteban R. G.; De Waele, Jo
Calculating flux to predict future cave radon concentrations, Rowberry, Matt; Marti, Xavi; Frontera, Carlos; Van De Wiel, Marco; Briestensky, Milos
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Your search for troglophile (Keyword) returned 35 results for the whole karstbase:
Showing 1 to 15 of 35
The birth of Biospeleology., 1964, Motas Constantin
Modern biospeleology dates from May 15, 1907, with the publication of Racovitza's "Essai sur les problmes biospologiques." In this paper he posed; if he did not answer; every question raised by life in the subterranean world. He outlined a program of biospeological research, made an analysis of the conditions of existence in the subterranean domain and their influence upon cavernicoles, discussed the evolution of subterranean biota, their geographical distribution, etc. Racovitza modified Schiner's (1854) classification, dividing cavernicoles into troglobites, troglophiles and trogloxenes, terms later adopted by a great number of biospeologists. The "Essai", called "Racovitza's famous manifest" by Vandel, was considered the birth certificate of biospeology by Antipa (1927) and by Jeannel (1948), its fundamental statute. Jeannel also made major contributions to the young science through his extensive and detailed studies. The names of Racovitza and Jeannel will always be linked as the uncontested masters of biospeology, the founders of Biospeologica, and the authors of Enumration des grottes visites. Apart from Schiner, whose ecological classification of cavernicoles was utilized and modified by Racovitza, they had another forerunner in Vir, a passionate speleologist who often accompanied Martel in his subterranean explorations, once meeting with a serious accident in which he was on the brink of death. Vir (1897, 1899) studied subterranean faunas, establishing the world's first underground laboratory, where he carried on unsuccessful or ill-interpreted experiments. We consider Racovitza and Jeannel's criticism of him too severe. Let us be more lenient with our forerunners, since their mistakes have also contributed to the progress of science, as well as exempting us from repeating them.

Remarks on the Japygidae (Insecta, Diplura) reported for the underground environment., 1964, Pages Jean
About 50 japygids, belonging to 29 distinct forms of which 23 are recognizable, have been collected since 1874 in caves all over the world. A list is given, by continent and by countries. Ten species found both in the soil and in caves are called troglophiles to emphasize the sorting which seems to occur among endogenous species. Of the remaining 13 species, all considered troglobites, only 3 show morphological peculiarities which can be ascribed to adaptation to cavernicolous life: (1) Metajapyx moroderi ssp. patrizianus Pags from Sardinia shows a slightly longer l0th urite and cerci than the f. typ.; (2) Kohjapyx lindbergi Pags from Afghanistan is characterized by its very long l0th urite, its relatively slender cerci, and the presence of more than 8 placoid sensillae (maximum basic number in endogenous species) on the apical segment of the antennae; (3) Austrjapyx leleupi Pags from the Lower Congo fits most closely the picture of the true troglobite; almost entirely depigmented, slender, with elongate legs, long setae, and the antennae with two of the trichobothria 4 to 5 times as long as the other typical 11, as well as 14 placoid sensillae on the apical segment. It is noted in the conclusion that, among the Diplura and Myriapoda, the almost exclusively phytophagous or saprophagous Campodeids and millipedes include a large number of true troglobites, in contrast with the carnivorous Japygids and centipedes, which have very few troglobites.

Contribution to the study of certain Lithobiidae (Chilopoda) of Romanian caves., 1965, Negrea Stefan
This paper comprises the ecological and zoogeographical data of 21 systematic units of Lithobiidae coming from 100 Romanian caves (Transylvania, Banat, Oltenie and Dobroudja). Initially the author describes in a summarising way the provenance of the studied material, after which he exposes fro every species the results of observations carried out by several authors and by himself. Finally he formulates the conclusions concerning ecology and biogeography resulting from his observations. These data are summarised in a synoptic table in which three species can be considered troglobionts, two of which are blind; these are Harpolithobius oltenicus Negrea; Lithobius decapolitus Matic, Negrea et Prunescu; Lithobius daeicus Matic; The other 18 species found are 5 troglophiles, 12 sub-troglophiles and 1 trogloxene.

Present-Day Cave Beetle Fauna in Australia A Pointer to Past Climatic Change, 1965, Moore, B. P.

Beetles form an important element of life in caves, where they provide some of the most spectacular examples of adaptation to the environment. The troglobic forms are of greatest interest from the zoogeographical point of view and their present distributions, which are largely limited to the temperate regions of the world, appear to have been determined by the glaciations and later climatic changes of the Quaternary. Troglophiles, which are much more widespread, show little adaptation and are almost certainly recently evolved cavernicoles.


Tachycines asynamorus Adel., eine troglophile Heuschreckenart., 1967, Lovcik, P.

Cockroaches (Blattodea) From Australian Caves, 1967, Richards, Aola M.

Ten species of Australian cockroaches are recorded from Australian caves and mines. Most are troglophiles or guanobia. Only one troglobitic species is known. The distribution of these species is given, and attention is drawn to their absence from south-eastern Australia and Tasmania. It is suggested that climatic changes in the Pleistocene and early Recent may have been responsible for this, and that the fauna found in many cave areas may be of comparatively recent origin.


Tasmanian Cave Fauna: Character and Distribution, 1967, Goede, A.

The geology and nature of the caves is discussed. Cave development has been affected by glacial outwash and periglacial conditions which must be taken into account when considering the development and distribution of cave fauna. The food supply in the caves is limited by the absence of cave-inhabiting bats. Floods while adding to the food supply must be destructive to some forms of terrestrial cave life. The cave fauna consists entirely of invertebrates. The carab genus Idacarabus Lea contains the only troglobites found in Tasmania. A common troglophile throughout the island is Hickmania troglodytes (Higgins and Petterd) which belongs to a very small group of relict spiders. Five species of cave crickets are known from Tasmania and Flinders Island. Three species belong to the genus Micropathus Richards and show an interesting distribution pattern. A single species of glow-worm, Arachnocampa (Arachnocampa) tasmaniensis Ferguson occurs in a number of Tasmanian caves. It is more closely related to the New Zealand species than to glow worms found on the Australian mainland. Other terrestrial cave life is briefly discussed. Aquatic cave life is poorly known. The syncarid Anaspides tasmaniae (Thomson) has been recorded from several caves. It differs from epigean forms in reduction of pigment.


Tachycines asynamorus Adel, eine troglophile Heuschreckenart, 1967, Lovcik, P.

Ecological studies in the Mamoth Cave System of Kentucky. I. The Biota., 1968, Barr Thomas C.
The Mammoth Cave system includes more than 175 kilometers of explored passages in Mammoth Cave National Park, Kentucky. Although biologists have explored the caves intermittently since 1822, the inventory of living organisms in the system is still incomplete. The present study lists approximately 200 species of animals, 67 species of algae, 27 species of fungi, and 7 species of twilight-zone bryophytes. The fauna is composed of 22% troglobites, 36% troglophiles, 22% trogloxenes, and 20% accidentals, and includes protozoans, sponges, triclads, nematodes, nematomorphs, rotifers, oligochaetes, gastropods, cladocerans, copepods, ostracods, isopods, amphipods, decapods, pseudoscorpions, opilionids, spiders, mites and ticks, tardigrades, millipedes, centipedes, collembolans, diplurans, thysanurans, cave crickets, hemipterans, psocids, moths, flies, fleas, beetles, fishes, amphibians, birds, and mammals. The Mammoth Cave community has evolved throughout the Pleistocene concomitantly with development of the cave system. The troglobitic fauna is derived from 4 sources: (1) troglobite speciation in situ in the system itself; (2) dispersal along a north Pennyroyal plateau corridor; (3) dispersal along a south Pennyroyal plateau corridor; and (4) dispersal across the southwest slope of the Cumberland saddle merokarst.

Ecological studies in the Mamoth Cave System of Kentucky. I. The Biota., 1968, Barr Thomas C.
The Mammoth Cave system includes more than 175 kilometers of explored passages in Mammoth Cave National Park, Kentucky. Although biologists have explored the caves intermittently since 1822, the inventory of living organisms in the system is still incomplete. The present study lists approximately 200 species of animals, 67 species of algae, 27 species of fungi, and 7 species of twilight-zone bryophytes. The fauna is composed of 22% troglobites, 36% troglophiles, 22% trogloxenes, and 20% accidentals, and includes protozoans, sponges, triclads, nematodes, nematomorphs, rotifers, oligochaetes, gastropods, cladocerans, copepods, ostracods, isopods, amphipods, decapods, pseudoscorpions, opilionids, spiders, mites and ticks, tardigrades, millipedes, centipedes, collembolans, diplurans, thysanurans, cave crickets, hemipterans, psocids, moths, flies, fleas, beetles, fishes, amphibians, birds, and mammals. The Mammoth Cave community has evolved throughout the Pleistocene concomitantly with development of the cave system. The troglobitic fauna is derived from 4 sources: (1) troglobite speciation in situ in the system itself; (2) dispersal along a north Pennyroyal plateau corridor; (3) dispersal along a south Pennyroyal plateau corridor; and (4) dispersal across the southwest slope of the Cumberland saddle merokarst.

The terrestrian Isopoda of the caves from Eregli to the Black Sea., 1971, Strouhal Hans
Dr Klaus Dobat (Tubingen) and Madame Dobat have collected the following terrestrial Isopods from caves of Eregli on the coasts of the Black Sea, North of Turkey: 1) Ligidium (Ligidium) assimile sp. nov., trogloxene widely distributed in the northern Anatolian territories; 2) Chasmatoniscus oculatus gen. et sp. nov., troglophile; 3) Trichoniscus (Trichoniscus) heracleotis sp. nov., troglobic; 4) Cylisticus (Platycylisticus) dobati subgen. et sp. nov., troglophile; 5) Cylisticus (Cylisticus) mechthildae sp. nov., troglobic. The author gives the descriptions together with figures of all these new Oniscoidea in this paper, M. Dobat describes the caves of Eregli and lists the animals herein found.

The terrestrian Isopoda of the caves from Eregli to the Black Sea., 1971, Strouhal Hans
Dr Klaus Dobat (Tubingen) and Madame Dobat have collected the following terrestrial Isopods from caves of Eregli on the coasts of the Black Sea, North of Turkey: 1) Ligidium (Ligidium) assimile sp. nov., trogloxene widely distributed in the northern Anatolian territories; 2) Chasmatoniscus oculatus gen. et sp. nov., troglophile; 3) Trichoniscus (Trichoniscus) heracleotis sp. nov., troglobic; 4) Cylisticus (Platycylisticus) dobati subgen. et sp. nov., troglophile; 5) Cylisticus (Cylisticus) mechthildae sp. nov., troglobic. The author gives the descriptions together with figures of all these new Oniscoidea in this paper, M. Dobat describes the caves of Eregli and lists the animals herein found.

The cavernicolous japygidae of the French fauna., 1972, Pages Jean
Twelve species of Japygids are cites from France. Six are known with certitude from caves. None of them shows any morphological modification which can be ascribed to cavernicolous life. All of them are found outside caves except Metaj. doderoi Silv. which is known from a unique female from Castel Mouly cave. To take into account the selection which seems to occur among the species, the term "subtroglophile" is proposed to include those which can be found in the caves.

The cavernicolous japygidae of the French fauna., 1972, Pages Jean
Twelve species of Japygids are cites from France. Six are known with certitude from caves. None of them shows any morphological modification which can be ascribed to cavernicolous life. All of them are found outside caves except Metaj. doderoi Silv. which is known from a unique female from Castel Mouly cave. To take into account the selection which seems to occur among the species, the term "subtroglophile" is proposed to include those which can be found in the caves.

A Check list of the Irish Cave Fauna: Troglobite, Troglophile and Trogloxene, 1974, Hazleton M.

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